The research reports on this page have been produced from a variety of sources including: undergraduate and graduate student researchers, research interns, and visiting scientists.

Contact with any questions regarding the research found on this page.


Esch, Clarice. 2013. Gunnera at Cloudbridge Nature Reserve. - Western Kentucky University

Mazzolari, Ana C.. 2008. Invasive Plant Species of Cloudbridge Reserve.

Medrano, Jacqueline Z.. 2008. Project Follow-Up Report: Sampling and Identification of the Flora of a Cloud Forest reserve in Costa Rica. - Arizona State University

Medrano, Jacqueline Z.. 2007.  Sampling and identification of the flora of a cloud forest reserve in Costa Rica. - Arizona State University

Valliere, Debbie L. and Dario A. Elliott. 2007. Determinates of Canopy Epiphyte Abundance in a Primary Lower Montane Cloud Forest in Cloudbridge Nature Reserve, Costa Rica.

Wolinsky, Cara. 2004. Flowering Bromeliads of the Chirripo National Park Region.


Frehner, Alena. 2018. The Influence of Habitat Factors on Species Richness and Abundance of Animals in a Montane Cloud Forest. - Van Hall Larenstein University of Applied Sciences

Vanderwalle, Cyrielle. 2017. Mammal and Sapling Trees Interactions. - AgroSup Dijon

Van Meter, Lara. 2016. An Analysis of the Invasive Potential of Cupressus lusitanica and Its Effects on the Chemical Properties of the Surrounding Soils. - University of Colorado at Boulder

Schoettlé, Guillaume. 2012. Study of the Distribution of Animal Species in a Rainforest. - AgroSup Dijon

Taylor, Joseph. 2007. Observations of hummingbird feeding behavior at flowers of Heliconia beckoner and H. tortuosa in southern Costa Rica. Ornitologica Neotropica 17: 133-138.

Taylor, Joseph. 2005. Do Sympatric Heliconias Attract the Same Species of Hummingbird? Observations on the Pollination Ecology of Heliconia beckneri and H. tortuosa at Cloudbridge Nature Reserve. - University of Glasgow


Laan, Chiel van der. 2018. Forest assessment of planted, naturally regenerated and primary tropical cloud forest. - Van Hall Larenstein University of Applied Sciences

Lefebvre, Cléa. 2018. Comparaison de peuplements issus de regeneration naturelle et de plantation. (Comparison of naturally regenerated and planted forest stands.) - Agro Paris Tech

Fleer, Michon. 2017. Reforestation practices at Cloudbridge Nature Reserve, Costa Rica. - Van Hall Larenstein University of Applied Sciences

Sanden, Mathijs van der. 2017. Habitat assessment of planted, naturally regenerated and primary tropical cloud forest. - Van Hall Larenstein University of Applied Sciences

Barz, Marian. 2016. Forest succession and fruit availability in planted and natural regenerated areas of Cloudbridge Nature Reserve, Costa Rica. - Van Hall Larenstein University of Applied Sciences

DeLyser, Kendall. 2015. Assessing the Effectiveness of Reforestation Efforts in the Tropical Montane Cloud Forest of Costa Rica.

Lehmann, Stephen. 2014. Untersuchung zur Eignung von Baumarten und waldbaulichen Verfahren für Restaurationsmaßnahmen im Nebelwaldgebiet Costa Ricas. (Evaluation of the Suitability of Certain Tree Species and Silvicultural Approaches for the Establishment of Restoration Areas of a Montane Oak Cloud Forest of Costa Rica). German with English (p.188) and Spanish (p. 200) summaries - Technische Universität Dresden (Dresden University of Technology)

Fröling, Marloes. 2012. Weed Suppression in Cloudbridge Nature Reserve, Costa Rica. - Van Hall Larenstein University of Applied Sciences

Spek, Michiel and Tijmen Hoogendijk. 2011. Cloud Forest Recovery - Evaluation at Cloudbridge Nature Reserve, Costa Rica. - Van Hall Larenstein University of Applied Sciences

Bol, Matthijs and Dennis Vroomen. 2008. The Succession of Pasture Land into Original Cloud Forest - In the pre-mountain area of Costa Rica.

Lehmann, Stephan. 2007. Tree Monitoring and Growth Analysis at Cloudbridge Nature Reserve (Forestry Economics and a Comparison of the Growth of Tropical Tree Species in the First Three Years of Growth). - University of Dresden

Isotupa, Olli. 2005. An Evaluation of Forestry Methods at Cloudbridge: Comparison with commonly used and recommended forestry methods in tropical forests, especially in Costa Rica.

Polach, Ivo and Marco A.O. Rojas. 2004. Monitoring and Assessment of Biodiversity at the Cloudbridge Nature Reserve, Costa Rica, Interim Project Report.
- Smithsonian Institution Monitoring and Assessment of Biodiversity Program & Herbarium of Universidad Nacional Heredia

Geology, Meteorology, & Hydrology Studies

Pfammatter, Prisca. 2017. Tropical montane cloud forest soils: A soil study in Cloudbridge Nature Reserve.

Whitley, Matthew. 2013. Cloudbridge Meteorological Update 2013.

Whitley, Matthew. 2013. Rainfall Variation Based on Forest Type.

Glide, Kailen. 2011. An Analysis of the Dissolved Oxygen and Volume Flow in Jilguero Stream, Cloudbridge Nature Reserve, Costa Rica.

Davis, Emily. 2009. Hydrogeology of an Upper Montane Tropical Forest. - SUNY New Paltz

Schembre, Charles. 2009. Soil Types and Fundamental Soil Properties of a Costa Rican Tropical Montane Cloud Forest.

Tingerthal, John, Sebastian Culbreth, Jessican Schwartz, and Tom Gode. 2007. GIS (Geographic Information Systems) Studies and Maps of Cloudbridge.

Giddy, Ian. 2006. Rainfall and Temperature.

Souers, Joanna. 2004. A Preliminary Study of Stream Chemistry on the Rio Chirripó Pacifico from the Chirripo National Park to Canaan, Costa Rica.

Land Use and Social Context

Gerolemou, Rosie. 2018. Reducing conflict between livestock owners and predators in Costa Rica. [Gente y carnívoros en Costa Rica.]

Rudman, Alexie, Emillee Hernandez, and Nainika Lamba. 2015. The Salstenible Project. - Borderless World Volunteers, McGill University

Marciano, Oren. 2010. Evaluación del Impacto del Ecoturismo Sobre las Actitudes y Creencias Relacionadas a la Vida Silvestre y, la Conservación de los Recursos Naturales, En Cuatro Comunidades Costarricenses. (Evaluation of the Impact of Ecotourism on the Attitudes and Beliefs Related to Wildlife and the Conservation of Natural Resources in Four Costa Rican Communities). - Universidad Nacional (National University)

Culbreth, Sebastian. 2007. Relationships Between Land Uses and Water Quality in Costa Rica: An analysis of the San Gerardo de Rivas watershed and the Chirripo River.

Culbreth, Sebastian Gonzalez and Tom Newman. 2007. Chirripó Valley Land Use Study – Watershed Analysis/ Análisis de Cuenca Hidrográfico, San Gerardo de Rivas, Perez Zeledon, Costa Rica. – A study facilitated by Cloudbridge Nature Reserve/ Una investigación facilitada por la Reserva Cloudbridge.

Stoll, Angelika. 2006. Sustainable Use of Forest Resources.

Schwartz, Jessica. 2004. Land Use at Cloudbridge and Vicinity.


Saunier, Baptiste. 2016. Etude Mycologique au Sein de la Foret de Cloudbridge (Costa Rica) (Mycology Study in the Cloudbridge Forest (Costa Rica)). - Université des Sciences et Techniques de Limoges (Science and Technical University of Limoges)

Carr, Reana. 2005. Mycorrhizal Density Differences in a Costa Rican Reforestation Project. - Cornell University

Zoology - Amphibians and Reptiles

Kauffmann, Emilie. 2015. Frog Survey During the Wet Season in Primary and Secondary Forests. - Engineering School of Agronomy and Food Process

van Kessel, Jasper. 2015. Herpetofauna survey secondary and primary forest. - Van Hall Larenstein University of Applied Sciences

Lau, Yan-Yee. 2012. Assessing the Species Composition Abundance of Anurans in Secondary Tropical Forests. - University of York

Cousineau, Susan. 2007. Cloudbridge Herpetofauna Survey: Part 1, Amphibians.

Jacobs, Toby. 2007. A Survey of Anuran Distribution in a Costa Rican Cloud Forest. - James Madison University

Michelsohn, Moses, Stephanie Sunderman, Pradip Ananda, Guy III (Trey) Kieckerhefer, and Christopher Wilkinson. 2007. Cloudbridge Herpetological Survey.

Zoology - Birds

Straub, Alyce. 2018. Resplendent Quetzal Literature Review

Cannon, Laura. 2017. The Effect of an Artificial Feeder on Hummingbird Behaviour. - University of Salford Manchester

Kane, Spencer, Neil Hancart, and Jennifer R. Powell. 2017. Bird Monitoring Study Data Report: January - February 2017.

Powell, Jennifer R. 2017. Bird monitoring study data report: Jan 2013 - Dec 2016.

Sutley, Amy and Baley Good. 2017. Bird monitoring study data report: July 3 - August 18, 2017. October.

Thierry, Mélanie. 2016. The Effectiveness of Tropical Cloud Forest Restoration on Bird Community at Cloudbridge Nature Reserve, Costa Rica.

Rabone, Anna and Christian Staunton. 2015. Hummingbird Behavioural Study at Cloudbridge Nature Reserve.

Stanley, Richard. 2012. Effects of Habitat Disturbance On A Montane Bird Community. - Harvard University

Dibala, Ryan. 2007. Avifaunal Diversity in Various Habitat Types in Costa Rican Montane Cloud Forest with the Implementation of CyberTracker.

Paradis, Eugene D.. 2007. A Survey of Tropical Owl Population Density and the Vocal Behavior of the Mottled Owl (Strix Virgata) In a Partially Fragmented Cloud Forest Habitat.

Marcy, Nathan.2007a. Cloudbridge Bird Diversity and Distribution: Mist Netting and Banding. July.

Marcy, Nathan.2007b. Bird Diversity at Cloudbridge: A Preliminary Report.

Marcy, Nathan. 2004. Bird Diversity and Distribution Avifauna Diversity at Cloudbridge.

Zoology - Invertebrates

Poznansky, Frederica and Jennifer R. Powell. 2017. Comparison of diversity, richness, and relative abundance of dung beetles in Cloudbridge Nature Reserve. (Spanish)

Memory, Andrew E.. 2009. Effects of Reforestation on Benthic Macroinvertebrate Diversity and Assemblage in Costa Rican Headwater Streams.

Peckmezian, Tina. 2009. A Baseline Study of the Spider Fauna at a Costa Rican Cloud Forest Reserve.

Henning, Bridget. 2008. Odonata of Cloudbridge: An inventory of dragonflies and damselflies in various habitats.

Rathbone, Sarah. 2007. The Lepidoptera Diversity of a Lower Mountain Cloud Forest In Costa Rica.

Vey, Aiden. 2007. Adaptations of Insects at Cloudbridge Nature Reserve.

Sheehan, Emily. 2007. Diversity and Abundance of Subsoil and Leaf Litter Invertebrates Across Different Levels of Disturbance in a Costa Rican Cloud Forest.

Brown, Alison. 2006. Effects of Habitat Alteration and Seasonal Variation on the Abundance and Diversity of Pterygota at Cloudbridge Nature Reserve, Costa Rica.

Rosser, Nick. 2006. Effects of Habitat Alteration on the Abundance and Diversity of Pterygota at Cloudbridge Nature Reserve, Costa Rica.

Zoology - Mammals

te Beek, Ramon. 2018. A habitat suitability study for sloth species Bradypus variegatus and Choloepus hoffmanni in Cloudbridge Nature Reserve, Costa Rica.

Meijboom, Wouter. 2013. High Altitude Cloud Forest: A Suitable Habitat for Sloths? - Van Hall Larenstein University of Applied Sciences

Cailly, Caroline, Benoit Duffay, and Margaux Mazille. 2010. Medium and Large Mammal Monitoring. (Suivi des Moyens et Grands Mammiferes.)

Marrocoli, Sergio. 2007. Small mammal assemblages at different levels of disturbance on the Cloudbridge Reserve.

More Resources

For more information on the local flora and fauna, check out our Fact Sheets.  Or check out the websites and books of some of our favorite organizations and authors.  If you are interested in learning which species can be found in Cloudbridge, take a look at our species lists.
Interested in becoming a Volunteer or Researcher?  Visit our Volunteer/Researcher Page to learn more, or fill out an application.


Comments are closed.